다양함이 가득한 #코리도라스 갈래 ( feat.corydoradinae Lineages on corydoras zon)
2021. 1. 5. 7:07
Corydoradinae Linages
다양한 코리도라스 갈래
A lot of change is coming to the naming and organisation of the Cory cats
Table of Contents
Chart of relation between Lineages
Map of Geographical distribution of mimetic communities
Lineage 1 - Saddle-nosed Corydoras
Lineage 9 - "classic short-snouts"
Overview
The Corydoradinae sub-family today (2020) consists of the following family names:
Aspidoras
Brochis
Corydoras
Scleromystax
The other sub-family is Callichtyinae, and they both belong to the family Callicthyidae.
In a paper "Evolution, Ecology and Taxonomy of the Corydoradinae Revisited", from 2011 by Markos A Alexandrou & Martin I Taylor a new systematic revision based on 9 Lineages was proposed. This website is using this article as a base for grouping the different species. They suggest new names or resurrected names for some of the lineages, but we have to wait for further research before they can be actually used.
It is however not as clear cut as that. One of the lineages, Lineage 8, has four sub groups, Sub-clade 1 (Brochis) through 4, which complicates things. Sub-clades 2 through 4 will need new family names. Also 2 species in Lineage 2 seems to be something other than Aspidoras and might also need a new family name. This means we might end up with the four families above split into 12 families!
Regardless it's clear that there will be huge changes coming to our Corydoradinae taxonomy over the coming years.
Chart of relation between Lineages
Original can be found here: ResearchGate
Mimetic species are different species looking alike, I will write an article later about this..
Phylogenetic relationships of Corydoradinae including co-mimics.
The pie chart shows percentage of mimetic species per lineage. Branches with mimetic species at tips are indicated with coloured circles (coded by lineage). Nodes with support below 0.8 (Bayesian inference; BI) probability and 70% (maximum likelihood; ML) are denoted with black open circles. Codes on pictures indicate snout types as determined by morphometrics and genetic lineage (L, long; S, short; IS, intermediate short; XL, extra long; IL, intermediate long). Representative images of morphotypes and colour patterns clockwise from lineage 1: (L-1) Corydoras maculifer, (L-1) C. simulatus, (L-1) C. sp. C109, (L-1) C. sp. C92, (L-1) C. narcissus; (S-2) A. poecilius*; (L-3) S. prionotus; (IS-4) C. mamore; (IS-5) C. sp. CW19, (IS-5) C. nijsseni; (S-6) C. paleatus, (S-6) C. nattereri; (S-7) C. sp. CW26; (XL-8) C. multiradiatus*; (IS-8) C. sodalis*; (IL-8) C. imitator, (IL-8) C. sp. CW6, (IL-8) C. seussi, (IL-8) C. sp. C122; (S-9) C. sp. C91, (S-9) C. gossei, (S-9) C. adolfoi, (S-9) C. metae, (S-9) C. araguaiaensis, (S-9) C. arcuatus, (S-9) C. julii. *Non-mimetic taxa.
Map of Geographical distribution of mimetic communities
Original can be found here ResearchGate.
Geographical distribution of mimetic communities
Genetic lineages are denoted by coloured circles; small grey rectangles represent independent mimetic communities numbered 1–24. Larger black rectangles indicate communities belonging to the same drainage or basin. Grey ellipses indicate approximate geographical distribution. Species images: (1) C. paleatus, C. ehrhardti; (2) C. nattereri, Scleromystax prionotus; (3) S. barbatus, S. macropterus; (4) C. maculifer, C. sp. C122, C. araguaiaensis; (5) C. julii, C. sp. C109; (6) C. oiapoquensis, C. condisciplus; (7) C. sp. C135, C. sp. C136; (8) C. evelynae, C. sp. CW13; (9) C. kanei, C. crimmeni; (10) C. sp. CW19, C. sp. CW26; (11) C. metae, C. simulatus; (12) C. imitator, C. adolfoi, C. nijsseni; (13) C. serratus, C. cf. arcuatus; (14) C. narcissus, C. sp. CW6, C. arcuatus; (15) C. sp. C84, C. sp. C156; (16) C. sp. CW28, C. pulcher; (17) C. trilineatus, C. leopardus; (18) C. tukano, C. sp. CW11; (19) C. sp. C91, C. sp. C92; (20) C. similis, C. sp. C66, C. ourastigma; (21) C. cruziensis, C. mamore; (22) C. gossei, C. seussi; (23) C. sterbai, C. haraldshultzi; (24) C. sp. C76, C. sp. C77.
The C- and CW-Number systems
The system with C-numbers was introduced in 1993 in the German Aquarium Magazine DATZ and covers the sub-family Corydoradinae of the Callichtyidae catfish.
When they started this numbering it was mainly used for new species that were collected but there were still a lot of species that were already established in the hobby that never got a C-number.
The CW-numbers from Ian Fuller's site CorydorasWorld.com is filling that gap and adding species that are not covered by the DATZ Magazine.
Here is a great article over on Scotcat.com that explains it in more detail: What the heck is a CW number?
Lineage 1 - Saddle-nosed Corydoras
This base group of "saddle-nosed" species should remain as Corydoras, as first described by Lacépède in 1803. The type species for the genus will remain Corydoras geoffroy. Long snouted species such as Corydoras fowleri are at the base of the Corydoradinae group.
Described species
Corydoras acutus, C. amapaensis, C. areio, C. aurofrenatus (CW005), C. blochi (C032), C.cervinus (C028), C. coriatae, C. cortesi, C. desana (CW011), C. ellisae, C.fowleri, C. geoffroy, C. heteromorphus, C. maculifer, C. narcissus, C. negro, C. orcesi, C. ourastigma, C. oxyrhynchus, C. pastazensis, C. saramaccensis, C. sarareensis, C. semiaquilus, C. septentrionalis, C. serratus, C. simulatus, C. solox, C. stenocephalus (C017), C. treitlii, C. vittatus, C. zawadzkii
C-numbers
(from Datz Magazine)
C008, C016, C017 = C. stenocephalus, C024, C028 = C.cervinus, C029, C032 = C. blochi, C038, C042, C047, C051, C053, C061, C063, C077, C078, C086, C092, C094, C095, C099, C109, C115 = C116, C116 = C115, C124, C127, C145, C146, C149, C153, C155 (either here in Linage 1 or in Lineage 8 Sub-clade 4)
CW-numbers
(from Ian Fuller's website Corydoras World)
CW005 = C. aurofrenatus, CW011 = C. desana, CW012, CW017, CW053, CW055, CW059, CW066, CW073, CW075, CW080, CW083, CW118, CW120, CW121, CW124, CW128, CW143, CW149, CW156, CW164
Lineage 2 - Aspidoras
This group should remain as Aspidoras (Ihering, 1907) with the type species: A. rochai. All known Aspidoras belong to this lineage apart from A. pauciradiatus. Both C. gladysae and C. petrarcini seem more closely related to Aspidoras, but they lack certain traits defining Aspidoras. They might therefore require a new generic name (Calvino and Alonso, 2009). Further studies need to be made to better understand the relationships of these species to Aspidoras.
Described species
Aspidoras albater, A. azaghal, A. belenos, A. brunneus, A. carvalhoi, A. depinnai, A. eurycephalus, A. fuscoguttatus, A. gabrieli, A. kiriri, A. lakoi, A. maculosus, A. marianae, A. menezesis, A. mephisto, A. microgaleus, A. poecilius (C158), A. psammatides, A. raimundi, A. rochai, A. spilotus (C125), A. taurus, A. velites, A. virgulatus
C-numbers
(from Datz Magazine)
C035, C036, C037, C118, C119, C125 = A. spilotus, C158 = A. poecilus
CW-numbers
(from Ian Fuller's website Corydoras World)
CW052, CW119, CW126, CW141
Lineage 3 - Scleromystax
All known Scleromystax belong to lineage 3 and this groups should remain as Scleromystax (Günther, 1864) with the designated type species: Scleromystax barbatus.
Described species
Scleromystax barbatus, S. kronei, S. lacerdai (C015), S. macropterus, S. prionotus, S. reisi, S. salmacis
C-numbers
(from Datz Magazine)
C015 = S. lacerdai, C112, C113
CW-numbers
(from Ian Fuller's website Corydoras World)
CW038, CW042, CW067, CW092, CW147, CW148
Lineage 4 - "Dwarf Species"
Lineage 4 includes two of the dwarf species, so they suggest to resurrect the disused Microcorydoras (Myers, 1953), with the designated type species: Corydoras hastatus. Oddly enough not all species in this group are real dwarfs, but they are closely related and have similar color patterns throughout the larval development.
Also note that C. habrosus is not included here.
Described species
Corydoras guapore, C. hastatus, C. mamore (C011), C. paucerna, C. pygmeaus
C-numbers
(from Datz Magazine)
C011 = C. mamore
CW-numbers
(from Ian Fuller's website Corydoras World)
CW153
Lineage 5 - "elegans" group
This group contains species known as the ‘elegans’group. During a revision the genus name Gastrodermus could be resurrected, with the designated type species Corydoras elegans. C. gracilisis is the basal species in this group and Aspidoras pauciradiatus belongs to this lineage rather than Aspidoras (Lineage 2)
Described species
Corydoras bilineatus, C. elegans, C. gracilis, C. nanus, C. napoensis, C. nijsseni (C111), C. sp. A. pauciradiatus, C. undulatus
C-numbers
(from Datz Magazine)
C041, C088, C089, C111 = C. nijsseni, C123, C126, C132
CW-numbers
(from Ian Fuller's website Corydoras World)
CW008, CW018, CW019, CW022, CW029, CW044, CW048, CW056, CW064, CW085, CW087, CW095, CW096, CW100, CW105, CW110, CW123, CW131, CW144
Lineage 6 - "true" Corydoras
Species within Lineage 6 have always been classified under the genus Corydoras. There are no synonymous disused generic names available so it would be necessary to describe a new genus with a new type species.
Described species
Corydoras albolineatus (previously misidentified as C. xinguensis, C056), C. carlae, C. cochui (C022), C. diphyes, C. ehrhardti, C. flaveolus, C: froehlichi, C. gryphus (C007 = CW024), C. lacrimostigmata , C. longipinnis (CW003), C. lymnades, C. micracanthus, C. nattereri, C. ortegai (CW031), C. paleatus, C. potaroensis, C. reynoldsi, C. steindachneri, C. tukano (sp. "Asher", C064)
C-numbers
(from Datz Magazine)
C007 = CW024 = C. gryphus, C022 = C. cochui, C040, C056 = C. albolineatus, C064 = C. tukano, C073, C114, C144
CW-numbers
(from Ian Fuller's website Corydoras World)
CW003 = C. longipinnis, CW024 = C007 = C. gryphus, CW031 = C. ortegai, CW077, CW108
Lineage 7 - "aeneus" group
This group contains all species within the ‘aeneus’ group. During a revision the genus name Osteogaster (Cope, 1871) would be resurrected , with the type species: Corydoras eques.
The most basal species in this group are Corydoras melanotaenia and C. aeneus from Trinidad. C. zygatus and C. rabauti are both within this group and seems to be more closely related to each other than they are to any other species in this group.
Most closely related to these species are C. aeneus spp. from the Parana drainage (originally known as C. macrosteus).
The Amazonian species form a group within this lineage, with C. aeneus from Suriname and Guyana separated from species from Peru where the ‘laser’ species are found.
Described species
Corydoras aeneus (spp.), C. eques, C. hephaestus, C. melanotaenia, C. rabauti, C. schultzei, C. venezuelanus, C. zygatus
CW-numbers
(from Ian Fuller's website Corydoras World)
CW007, CW009, CW010, CW014 (= CW023), CW016, CW023 (=CW014), CW026, CW041, CW043 (= C. eques), CW068, CW078, CW084, CW093, CW097, CW139
Lineage 8 - Brochis et. al.
Lineage 8 contains mainly the long snouted but deep bodied species, but also includes Brochis, that was recently synonymized with Corydoras (Britto, 2003). The name Brochis would most likely be resurrection during a revision, with the designated type species Brochis splendens. There are also 3 more sub-clades that would need new names.
Sub-clade 1 (remains Brochis)
Described species - Brochis
Brochis britskii, B. multiradiatus, B. splendens
CW-numbers
(from Ian Fuller's website Corydoras World)
CW034, CW035, CW132, CW136
Sub-clade 2 (new name needed)
Described species
Cordydoras costai, C. difluviatilis, C. filamentosus, C. garbei
C-numbers
(from Datz Magazine)
C057, C067
Sub-clade 3 (new name needed)
Described species
Corydoras geryi, C. latus, C. pantanalensis (C005), C. reticulatus, C. sodalis
C-numbers
(from Datz Magazine)
C005 = C. pantanalensis, C081
CW-numbers
(from Ian Fuller's website Corydoras World)
CW033, CW061
Sub-clade 4 (new name needed)
Described species
Corydoras agassizii, C. amandajanea (very similar to C. crypticus), C. ambiacus, C. approuaguensis, C. arcuatus (CW036), C. bifasciatus, C. brittoi, C. condisciplus, C. crimmeni, C. crypticus (very similar to C. amandajanea), C. delphax, C. ephippifer, C. gomezi, C. haraldschultzi, C. imitator, C. incolicana (C001), C. isbrueckeri (C050), C. lamberti, C. leopardus, C. noelkempffi (C058), C. ornatus, C. orphnopterus, C. pinheiroi, C. pulcher, C. robinae, C. robustus, C. seussi (C027), C. spectabilis, C. spilurus, C. sychri, C. virginiae (sp. "Miguelito", C004)
C-numbers
(from Datz Magazine)
C001 = C. incolicana, C004 = C. virginiae, C009, C010 = C080, C013, C018, C027 = C. seussi, C034, C039, C049, C50 = C. isbrueckeri, C052, C058 = C. noelkempffi, C066, C068, C071, C074, C075, C080 = C010, C087, C097, C098, C101, C102, C103, C110, C117, C122, C128, C130, C131, C135, C138, C140, C141, C143, C152, C155 (either here in Linage 8 or in Lineage 1), C156, C157, C159
CW-numbers
(from Ian Fuller's website Corydoras World)
CW002, CW006, CW013, CW020, CW025, CW040, CW057, CW058, CW065, CW070, CW072, CW074, CW082, CW089, CW090, CW099, CW101, CW106, CW113, CW116, CW117, CW130, CW134, CW135, CW138, CW150, CW151, CW155, CW157, CW160, CW161
Lineage 9 - "classic short-snouts"
The species in this lineage are the classic ‘short snouted’ species such as Corydoras adolfoi. When revisied it would likely mean the resurrection of the name Hoplosoma (Agassiz, 1846). The designated type species would be Corydoras punctatus. This lineage is very species rich and many of the them are have evolved relatively recently.
Described species
Corydoras acrensis, C. adolfoi, C. amphibelus, C. apiaka, C. araguaiaensis, C. armatus, C. atropersonatus, C. axelrodi, C. baderi, C. benattii, C. bicolor, C. boehlkei, C. boesemani, C. bondi (C031), C. breei, C. brevirostris, C. burgessi, C. caudimaculatus, C. concolor, C. copei, C. coppenamensis, C. cruziensis (C012), C. davidsandsi, C. duplicareus, C. esperanzae, C. evelynae, C. eversi (C065), C. gossei, C. granti (previously misnamed C. arcuatus; C020), C. griseus, C. guyanensis, C. habrosus, C. julii, C. kanei (C026, C046), C. knaackii, C. leucomelas, C. loretoensis, C. loxozonus (C082), C. melanistius, C. melini, C. metae, C. multimaculatus, C. oiapoquensis, C. osteocarus, C. panda, C. paragua, C. parallelus (C002), C. pavanelliae, C. polystictus, C. punctatus, C. rikbaktsa, C. sanchesi, C. schwartzi, C. similis, C. sipalwini, C. sterbai, C. surinamensis, C. trilineatus, C. urucu, C. weitzmani, C. xinguensis (C055, C105, C106, C107, C108)
C-numbers
(from Datz Magazine)
C002 = C. parallelus, C003 (sp. "Deckeri"), C006, C012 = C. cruziensis, C014, C019 = cf. evelynae, C020 (= C. granti), C021, C026 = C. kanei , C030, C031 = C. bondi, C033, C043 = C044, C044 (juvenile C043), C045, C046 = C. kanei , C048, C054, C055 = C. xinguensis, C062, C065 (C. eversi), C076, C079, C082= = C. loxozonus, C083, C084, C085 ("Mega metae"), C090, C091, C096, C100, C104, C105, C106, C107, C108 = C. xinguensis, C120, C121, C129, C133, C134, C136, C137, C139, C142, C147, C148, C150, C151, C154
CW-numbers
(from Ian Fuller's website Corydoras World)
CW001, CW004, CW015, CW021, CW027, CW028, CW030, CW031 (C. ortegai), CW032 (C. knaacki), CW036 (the true C. arcuatus), CW037, CW039,CW045,CW046, CW047, CW049, CW050, CW051, CW054, CW060, CW062, CW063, CW069, CW071, CW076, CW079, CW081, CW086, CW088, CW091, CW094, CW098, CW111, CW112, CW114 (Corydoras sipaliwini), CW115, CW122, CW125, CW127, CW129, CW133, CW137, CW140, CW142, CW145, CW146, CW151, CW152, CW158, CW159, CW162, CW163, CW165