Asperula aristata subsp. scabra (J. Presl & C. Presl) Nyman

Posted byTrajche DimitrovskiPosted inPlant familiesTags:,

Taxonomy: Fam. Rubiaceae// Genus: Asperula// Species: Asperula aristata L. fil.// Subspecies: Asperula aristata subsp. scabra (J. Presl & C. Presl) Nyman.

Determined according to the botanical key provided by Ehrendorfer and Krendl in Tutin et al. (1976).

Distribution: Native to Europe – France and South Europe to the Balkan Peninsula and Romania according to the distribution information provided by The Euro+Med PlantBase (source: Marhold, 2011+).

Habitat:  Open and dry habitats – grasslands, pastures, shrublands, rocks and rocky terrains.

Lifeform: Perennial, hemicryptophyte according to the Raunkiaer system.

Flowering period: Summer – autumn: JUN, JUL, AUG, SEP.

Asperula aristata subsp. scabra – habit in habitat. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – habit. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – habit. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – habit. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – habit. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – habit. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowering stem with spicate partial inflorescences. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowering stem with spicate partial inflorescences. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowering stem with spicate partial inflorescences. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – spicate partial inflorescence. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – spicate partial inflorescences. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – partial inflorescence with flowers. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowers. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowers. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – partial inflorescence with flowers. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowers. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowers and fruits. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowers and fruits. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – stem with flowers. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – stem with leaves. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – stem with leaves. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – stem with leaves. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – stem and leaves. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – stem with leaves. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – stems with leaves. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – stem and adaxial side of leaves. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – lower part of stems and plant base. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – lower part of stems and plant base. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – plant base and rootstock. Osogovo, mountain ridge between the gorges of “Kochanska” River and “Orizarska” River, 26.06.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – habit. Hilly region near village of Beli, 23.07.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – habit. Hilly region near village of Beli, 23.07.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowering stem with spicate partial inflorescences. Hilly region near village of Beli, 23.07.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowering stem with spicate partial inflorescences. Hilly region near village of Beli, 23.07.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowers. Hilly region near village of Beli, 23.07.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowers and fruits. Hilly region near village of Beli, 23.07.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowers and fruits. Hilly region near village of Beli, 23.07.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – stems with leaves. Hilly region near village of Beli, 23.07.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – stem with leaves. Hilly region near village of Beli, 23.07.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – stem with leaves. Hilly region near village of Beli, 23.07.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – lower part of stem. Hilly region near village of Beli, 23.07.2022. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – habit in habitat. Osogovo Mountains – “Mala” River, near lake “Gradche”, 27.06.2020. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – habit. Osogovo Mountains – “Mala” River, near lake “Gradche”, 27.06.2020. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – habit. Hills near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra (habit). Hills near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – flowering stems. Hills near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – spicate partial inflorescences. Hills near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – spicate partial inflorescences. Hills near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.
Asperula aristata subsp. scabra – lower part of stems. Hills near Kochani, 05.07.2020. Photo by Trajche Dimitrovski.

References:

Asperula aristata subsp. scabra. Marhold, K. (2011+): Rubiaceae (pro parte majore). – In: Euro+Med Plantbase – the information resource for Euro-Mediterranean plant diversity. Published on the Internet https://europlusmed.org/cdm_dataportal/taxon/0e996707-85ea-4604-95a2-51a52d2e8076 [Accessed 30.05.2022].

Asperula L. Ehrendorfer, F. and Krendl, F. in Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (1976). Flora Europaea. Volume 4: Plantaginaceae to Compositae (and Rubiaceae). Cambridge: Cambridge University Press, pp.4-14.

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